And what that has to do with bee husbandry?
You should trust that natural selection takes its job on British Isles by itself. Over 90% out of their beekeepers are 2 hive owners and they cannot fight against natural selection. They do not need any theories, what will happen in the future. The whole England is full of pure Blenheim bees.
Ok, I've understood you now, apologies. Most UK bees are kept by small beekeepers. And for them a measure of genetic control is hard to impossible.
First: its not natural selection they have to fight against! Natural selection for the FITTEST strains is what they need. And they do that best if left alone.
Second: I disagree - what UK beekeepers do need is an understanding of plain genetic husbandry.
I thought you denied the possibility of 'Blenheim bees'?
To your main point: yes, being small is a problem - for some. However:
1) there are a surprising number of areas where feral bees have re-established, having developed resistance through natural selection. If you are lucky enough to be able to keep bees near them you will likely have no problem, no matter how few hives you have. Just don't go buying packages or bringing duff queens in!
2) try to form a non-medicating club with nearby beekeepers.
3) locate as far as you can from large numbers of medicating beekeepers
4) try to mate your queens in promising feral areas
5) keep more than 2 hives!
6) when you get a promising colony, give the queen plenty of room. She'll raise more drones likely carrying good genes. And use her to requeen other hives.
7) imo its worth bearing in mind that having the right mite strains is also part of the solution. Given time the bees will probably raise them
So: yes, being in the wrong place makes it impossible to go treatment free. And being in a non-ideal place and having too few colonies makes it an uphill struggle. But most people will be able to evaluate their chances and form a plan.