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I have nursed hives without excluder the whole my life.
I want that queen may lay as much it can.
When I give to hives 3 brood boxes, queen does not need to rise to lay up to supers. When in main yield entrance is widely open, queen moves from lowest box because it is too cold, but bees fill it with pollen.
Lowest frames should be a little bit brood. Bees do not like to store pollen in white combs.
It depends on quality of queen, how much it is able to lay. Is one or two boxes enough to brood.
If ventilation is too big, queen may move even to the top most box to lay.
So open mesh floor will be the reason if the brood moves too high.
When I use electrict floor heating in spring, queen love to layn in lovest box.
When bees have good room to pollen near brood, they do not store it into supers.
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These are the basics how to keep brood, pollen and honey apart from each others.
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