MuswellMetro
Queen Bee
I read this a while ago which suggests that the population of the hornet is severely lacking in genetic diversity resulting in the early production of diploid drones which hinders the colony development.
In bees diploid drones aren't viable, I assume it is the same in hornets? If not I'm sure someone will tell me. I'm not sure if the hornets eat them like bees do.
Have you seen anything to suggest inbreeding is causing the hornets issues?
http://journals.plos.org/plosone/article?id=10.1371/journal.pone.0136680
this similar article to your link suggest they reach adulthood as diploid male adult hornets were caughtand also unpublished reference to triploid males ( queen mated with diploid drone)
https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4586377/
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