Take these scenarios
scenario 1 brood box with stores - bees will move en masse around the frames in the brood following the food - they will stay in their cozy little cluster with queen safely tucked nice and warm in the middle - bees will feed her.
scenario 2- brood box then super with stores -bees will move en masse around the frames in the brood following the food, bees will move up onto the frames in the super when food supply in the brood runs out - they will stay in their cozy little cluster with queen safely tucked nice and warm in the middle - bees will feed her.
scenario 3 - brood box, crown board with hole in, lump of fondant over hole - super as an eke over fondant -bees will move en masse around the frames in the brood following the food, when food runs out cluster will move until directly under hole and some bees will pop up, get fondant and tuck back into cluster - queen safely tucked up etc etc
scenario 4 - brood box, queen excluder then super with stores -bees will move en masse around the frames in the brood following the food with queen safely tucked, etc.etc, until stores run out in brood
then
either bees will move up onto the frames in the super queen left behind under QX
not safely tucked up etc etc therefore queen is dead
OR
Bees being bees refuse to abandon the queen to her fate thus all stay beneath the QX within sight but not reach of plentiful stores thus whole colony dies - ISOLATION STARVATION.
I know someone who this has happened to - not a beginner but an experienced beek (I won't embarrass the SBI in question by saying any more