Morning philip I can't explain why the colony have been in the bottom box all winter there is no barrier for them to cross,
When I removed the top box there was only three bees in it and the colony were covering 5/6 seams of bee's in the bottom.
In the autumn they were covering 7 in both.
It was the best thing to do removing the top box.
Looking at our records this colony did much the same last winter. They supersede in August to.
They also end up on double brood with out fail and are one of our biggest Amm colonys.
As to winter prep they were very light going into winter they were fed 16kgs of fondant.
My thoughts are they would benefit from being housed in a bigger hive type but then condensed into one for winter.
They were to big in the autumn to condence into one nats brood box.