I agree with all that.
Bees head down can be trying to keep warm as well.
There are patches that look like salt granules sprinkled about (again, photo is not the best way to tell) which are varroa faeces.
Best of luck......
I agree too, i would say sick bees trying to go in a long winter that was just too damn warm in the autumn.
As many have discussed and i hold up my hand and say i too, waited for a "brood break" and i know my Varroa levels were high, I didn't treat until Late December with vaped oxalic acid.
Big mistake, we all know that bees that are made in the autumn, are the ones that will live until right now and in to April. If these are sick before going in to winter the colony has to struggle right from the start of autumn.
I lost a few like this, with lots a tiny salt like deposits in cells where the brood nest was and bees then died . like starvation , but not all at once. Also a few individual brood cells that didn't hatch out because the colony wasn't able to heat this last emerging brood, as the colony finally collapsed.
Couple this with the long wet mild winter, that for us, terminated in a fairly chilly February and March its not the best scenario.
This year i am treating after that harvest and again in October, then again in mid winter. I feel i have to get numbers down early and winter is more of a mopping up job. Well thats what i feel might have happened.
Just dont beat yourself up about it, its a right pain but you will have better bees this year for sure! Onwards and upwards.