Itma,
Any space within a timber construction, with stagnant air, will likely become damp and be subject to mould and rot.
Your house roof must have ventilation to avoid rotting roof timbers, under-floor airbricks/grills must be maintained with houses with downstairs floorbooards, or the floor will rot (dry- or wet-rot). Cavity walls had air bricks installed for the same reason - there must be a small rate of air change to avoid damp conditions prevailing within the fabric of the structure.
My hive roofs are all ventilated to the exterior; it's just that my crownboards are not. That would be akin to leaving the attic door open all year round, or opening windows while using air conditioning or heating in the room! No timber product has a moisture of zero and if that moisture (and any other stray dampness) is all moved to one area, or allowed to accumulate, damp, mould and rot may well result. Ventilating the roof space is not the same as ventilating the hive into/through that space.
With an OMF there is more than adequate ventilation from below, for the cavity which is occupied by the bees, while air currents are not set up around/through the hive by convection any more than the bees require. Bees are perfectly able to maintain their space with their own air-con system when they are operational. They would have typically lived in any cavity in a tree, irrespective of whether there was more than adequate free air movement - as they can ventilate their space and control their hive temperature to quite close limits. They would not typically select a location, for setting up home, with a continual strong through-draught or one without protection from above.
Their comb structure would normally regulate any free air movement, fixed to the walls, ceiling and folding around the living area. We give them regimented parallel frames and try to avoid them making comb in other than the designated frames! We give them a hive with a designed wall thickness of 19mm (3/4 inch) or thereabouts.
Yes, we coddle them and provide extra food in the winter should they require it. In nature a lot of colonies would perish if they were only provisioned with around the 'bare minimum' of stores.
My polystyrene hives have a solid roof, with no holes in it. The bees are warm and provide their own air-con throughout the summer season. The time while they are clustered and inactive is the time when damp conditions must be avoided and ventilation must be adequate to maintain a dry living area for the bees. It only needs to be enough to clear about 15kgs of water over a 3 or 4 month period. Internal wall temperatures will be different with these different materials. 40mm of EPS has a far higher insulation value than any 19mm timber plank or a 5mm ply-board roof.
My crownboards are 9-10mm thick ply, btw. They have been treated and the inner sides often have a layer of wax, where comb has been built up to it and subsequently scraped away by me. There will not be much moisture penetration and the small amount of ventilation for the roof space has been adequate thus far (the crownboards are dry all through the winter). Condensation problems do not occur within the hive cavity because the temperatures are slightly elevated from the outside and that moisture, produced by the bees' respiration, has ample opportunity to diffuse/disperse through the OMF. Honey frames (and pollen alike), do not get moulded (as I found them when I started beekeping - when my mentor simply said 'the bees will clean the frames up shortly').
There is more than adequate ventilation through the OMF for every one of my colonies in my conditions, I can assure you of that. I recognise they will consume some extra stores with a gaping hole beneath them but that is acceptable - as long as the heat is not simply transferred quickly to the outside and lost.
Remember, all the heat we provide from our central heating system (or otherwise) in our house is lost to the surroundings over a very short time scale; every bit of fuel burned eventually goes towards heating up the outside environment - we just 'insulate and draught-proof' to keep it in our houses for a few extra hours.
Regards, RAB