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suggesting? thinking out loud, by what I can see from a quick look at the article, he doesn't say much about bigger being better, in fat at one point he sdays that small queen are as good, he talks more about fecundity and egg laying capabilities rather than size
Perhaps in one of the other articles in the series, he talks about larger queens being produced from larger eggs and the larger queens being more fecund because there is more space in their abdomen for storing sperm and producing eggs.
James