Hi BCrazy,
sorry, I must have been fast asleep. My understanding is that the 'organ of Johnston' detects the deflection of the flagellum by using a ring of sensory cells connected to the intersegmental membrane 'above' the pedicel (Anatomy and dissection of the honeybee, H A Dade. pp85). The deflection is translated into wind speed and direction and thus can 'tell' the bee the speed and direction of the airflow over it's antenna. Cross referenced with their known track (Direction in relation to the sun), it gives the bee an understanding of the speed and direction of the air mass in which it flies. While not wishing to bore you on the beautiful art of navigation, cross track error is the effect of a wind or tide on a steered course. In other words, the bees detect the difference between the direction they point when they fly (heading) and their real movement across the earth (track). The difference between the two is known as cross track error.
It’s important to bees because it’s information that needs to be added to the algorithm that makes up the waggle dance. No use telling your mates to fly at such and such angle to the sun, if the 10 knot wind from their right sends ‘em two miles downwind from the target.
I’ll try to find a source for this use of the OoJ, I think I may need to look at waggle dance references rather than the anatomical ones I’ve tried so far.
Rgds